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Rémy is a rat living in Paris who wants to be a chef, but has to overcome the disapproval of his family and the prejudice of humans. Using a somewhat hapless young man as a means through which to practice his art, Rémy meets yet another obstacle in the form of a famous, notoriously hard-to-please food critic named Anton Ego, nicknamed "The Grim Eater". The questions remain: Will Remy achieve his goal? Will he be able to please everyone in his locality?